The oldest known record of amphibian tracks in the UK has been discovered in a sandstone fossil.
New research has revealed an ancient trackway, found imprinted on a block of sandstone from the base of Hardraw Force Waterfall, in Wensleydale, North Yorkshire.
Dating back 340 million years, it is the oldest record of amphibian tracks in the UK.
The tracks belong to the earliest relatives of modern amphibians, called temnospondyls, specifically the edopoids, or "glutton-faced animals".
In order to see it in further detail, the trace fossil, currently on display at the Natural History Museum, was 3D scanned.
This was done as part of a research project by a previous undergraduate student from the University of Birmingham, Hannah Bird, who said: "We used scanning and photography to make a 3D digital model, allowing us to better visualise and identify the footprints and invertebrate traces."