Geraint Thomas has lost the leader's jersey in the Tour de Romandie after a puncture late on the second stage cost him over four minutes.
The 28-year-old punctured on the final climb of the stage into Saint-Imier.
Fellow Team Sky rider Chris Froome is now third in the overall standings after finishing in a reduced peloton.
Froome, who also punctured but recovered, is 10 seconds behind Michael Albasini who took Wednesday's 168.1km stage and with it the overall lead.
Tour de Romandie Stage 2 results
1. Michael Albasini (Switzerland / Orica) 4:21:43"
2. Jarlinson Pantano (Colombia / IAM Cycling) ST
3. Julian Alaphilippe (France / Etixx - Quick-Step)
4. Nathan Haas (Australia / Cannondale)
5. Rui Costa (Portugal / Lampre)
6. Damiano Caruso (Italy / BMC Racing)
7. Ilnur Zakarin (Russia / Katusha)
8. Ivan Santaromita (Italy / Orica)
9. Jan Bakelants (Belgium / AG2R)
10. Ramunas Navardauskas (Lithuania / Cannondale)
23. Chris Froome (Britain / Team Sky)
Overall classification after Stage 2
1. Michael Albasini (Switzerland / Orica) 4:42:52"
2. Ivan Santaromita (Italy / Orica) +10"
3. Chris Froome (Britain / Team Sky)
4. Simon Yates (Britain / Orica)
5. Ilnur Zakarin (Russia / Katusha) +15"
6. Pavel Kochetkov (Russia / Katusha)
7. Egor Silin (Russia / Katusha)
8. Yury Trofimov (Russia / Katusha)
9. Simon Spilak (Slovenia / Katusha)
10. Julian Alaphilippe (France / Etixx - Quick-Step) +20"